&L
I0(2
#)
P
%
'
I-/-!`:G:_9CBD>4)U>D:9Ga7d567;
7?>1@4 C>6>@D>E8"5"74B"#>9C<=7D"6O#C:
@>4
'3 %)
&
_=>@O">9:6404>9BdD>EG?#8"?7?"#@4?"7
B7 "#" "#O">9:B6>#A>B9C<?
51 trang |
Chia sẻ: huyen82 | Lượt xem: 2032 | Lượt tải: 0
Tóm tắt tài liệu Nghiên cứu sự phân hủy Lignin của một số nấm đảm và khả năng ứng dụng, để xem tài liệu hoàn chỉnh bạn click vào nút DOWNLOAD ở trên
$c;"
A 8 >" ;"! 9 >E6 CC ?: ]^ ;" BCG $c ;" A" >E9C
B9C ?AAA" 84" >E9C A ; 9 ?C ?#>
A?" 9>E6?:#4.#>F:9
P7I-/0!345G>>?`8?E@MDDYCO:73 %)
GU:7 !345G>>?`8?
E@>?6D96>?3<C&
!345G>>?`8?
E@>?6D96>?3<C;
,">C>97?#DOB0
_%
,"BCJ]^K
$^GS%
''Gc%
S%
!/
_]
_5
!K
!L
_c
_^
_L
_$%
_$'
_$]
_$S
_$c
_$^
_$L
_'%
_'$
_''
_']
_'5
_'&
_'S
_'c
_]&
_5]
_5&
_5S
_5^
!KW
!K/
_S$
_c%
_c$
_c]
_c5
_c&
_cS
_cc
,"AJ$cK
!I!L/
_^X_S'
/R^KW
$&GS%
$LG'%
/J^RW
V^WW
$LG%%
$&G5%
$$G&%
^G]%
cG'%
$%G$%
$&G$%
''G%%
$]G%%
$5G]%
cG]%
LG&%
$]G%%
$5G&%
$%G&%
$5G^%
$'G&%
$'GS%
$SG]%
$%Gc%
$'GS%
$]G$%
$%G&%
//^HW
/H^KW
$'G5%
'$G'%
$$G%%
$$G^%
^G&%
^G]%
$%G&%
LG&%
S^IW
$$G'%
IH^WW
$%G%%
'5G&%
HK^SW
HH^/W
''G&%
'$G&%
$5G5%
^G%%
LG%%
$&G'%
$cG^%
'$G&%
$LG&%
$]G^%
LG$%
$cG%%
$cG]%
$&G]%
$5G]%
$SG&%
$5G'%
$&G5%
$&G5%
$5G]%
$]GS%
$&G'%
$]G^%
HJ^WW
HI^KW
$$G&%
'5G%%
$'G^%
$$Gc%
$5G]%
$$G'%
$'G^%
$%Gc%
HI^WW
$cG&%
S$
_^X_S$
_^X_&5
_&X_&%
_]X_S%
_$X_^
_$X_c
_$X_&
_$X_]
_$X_$'
_$X_$5
_]X_$%
_&X_$%
_5X_S
_^X_S%
_SX_L
$]G%%
$$G%%
'$G%%
$^G%%
$LG'%
$^G%%
$LG5%
$SG%%
$cG%%
$]G%%
$%G'%
$$G]%
$5G5%
$%Gc%
$^Gc%
'%G&%
$LG&%
'5GL%
$cG]%
'5G&%
'%G%%
']G&%
$^G&%
$^G^%
LG5%
$$Gc%
LG'%
$&G%%
$$G'%
'%G&%
_9 >E6 CC ?" D7D B7 7 =: 3 >3 >E9C: ?: ?7?
?:G;"l?"D7D?6B"7?7?;">E68"E"?97
Z"!E>9#0;">":E89C>E"@C;""#?OB0
@I-/0+9C=<!I;\<G_9C=<!L/
S'
7 d 5 6 7 : ; ; ;9 8 \ C ; : 7
: 7 ! : Z 9 5 ; ; D : 9 D D
0
5
1 0
1 5
2 0
2 5
S C 3 S C 6 1 S C 3 - S C 6 1
7
D D Y C
:
: A
9>I-/07d567:;;;98\C;:7BD3 %)
7??:
D7>>E6>"#>>E?:4">E9C
A8G>OQD9CD7DAG>?"7>6?"<A"<?B9C<?9!
;"DC6>#;"A_]X_S$DC<D
8C74 D : ?AAA" G ?" ;" A _$X
_$5>H9C<?A<
,"BC_S?"7?0;D7>>E6 CCJ44bK>E4">E9C
qLG%%8 >E4" >E9C Aq''G$% A4"@>?: >"#> ?"7 >6 9:;<
?"?7?>1@4>#D>"
GD9CD7DA8V>9CB9C<?;
>E"?: >9C?"7 97? "#4<C;"BC
E"
F7 >EHA>6 >@B9C14 >
4"4"#GC:BAD:A>"#>81>D:?AAA" ?!?a
A"0A@?"?"
D:>#D>";"!F0>#@
' 2..% ).
S]
99:G=#>F: >@B9C<?$^;"
A.:3D7>>E6>E4">E9CA8?:]c;"BC8$^
;"A>6@C:O:]'
P7I-H0!345G>>?`8?E@MDDYCO:72..% ).
GU:7 !345G>>?`8?
E@>?6D96>?3<C&
!345G>>?`8?
E@>?6D96>?3<C;
,">C>97?#DOB0
%
,"BCJ]cK
$
H
]
5
&
S
c
^
L
$%
$$
$'
$]
$5
$&
$S
$c
$^
$L
'%
'$
''
']
HJ
HK
'S
'c
'^
'L
]%
]$
cG]%
'GS%
J^VW
'G$%
'GS%
'G5%
'Gc%
5G^%
^GS%
^GS%
5G]%
LG5%
SGS%
&Gc%
$]Gc%
&G%%
'G&%
5Gc%
5G'%
'G'%
&GS%
&G'%
5G'%
'G]%
/^LW
H^IW
$G]%
$G$%
]G'%
'GS%
'G^%
5G&%
LGc%
]G5%
S^/W
$G^%
&G&%
]G^%
'G&%
&GS%
'GS%
SGc%
5G$%
^G^%
cG'%
5G$%
&GS%
&G$%
]Gc%
]Gc%
'G$%
'G^%
5G]%
]G5%
]G^%
$G]%
L^RW
K^JW
'G$%
]Gc%
&GS%
'G%%
$Gc%
$G%%
S5
]'
]]
]5
]&
]S
IR
,"AJ$^K
$2'
'2c
52^
&2$5
^2L
^2$$
L2$'
$%2$5
$$2$'
$$2$]
$c2$^
$c2'$
$^2'&
$L2'$
'$2''
'$2'&
'$2'^
$%2'^
&G'%
'G]%
&G$%
SG%%
SG'%
K^LW
X
&G$%
&GS%
X
SG&%
&G^%
$%G$%
LG5%
SG'%
SG5%
SGc%
^G$%
SG5%
SG]%
^G'%
SGS%
cG]%
LG$%
]G&%
5G$%
SG&%
&G'%
^G&%
//^LW
LG&%
^G$%
]G%%
cG^%
^G]%
cG%%
X
$%GS%
^GS%
^G$%
&G%%
SG%%
cG'%
SG&%
^G^%
LG%%
$%G%%
X
)p4"@> "#;"BCI#>@=:3D:A>"#>81>D:
?AAA" 9'G'5G'&G]c
E" "#$^;"A?9>#I#>
@=:3G>#>#>1@4>E0>"6C<DACG
?"7>6>>@DB9CE
''4
.:" 7>=:3D: A >E$$;"BC85;" AG
@B9CF:9
P7I-I0!345G>>?`8?E@MDDYCO:74
S&
GU:7 !345G>>?`8?E@
>?6D96>?3<C&
!345G>>?`8?E@
>?6D96>?3<C;
,"BCJ$$K
NNgX
$
NNgX
&
NNgX
c
NNgX$'
NNgX$&
NNgXf'^
NNgX'$%
NNgX'$]
NNgX`$
NNgX`&
NNgX`S
,"AJ5K
NNgX`&XNNgX'$%
%%PL%%2K
%%/H%%eHS
NNgX$'XNNgX
c
&G]%
5G]%
&G&%
SG5%
5G&%
&G%%
5G&%
]G&%
&G%%
5G^%
&G'%
]G^%
H^HW
5G&%
&G^%
5G'%
$GS%
5G]%
5G5%
]G5%
'Gc%
$G$%
$GL%
]GS%
]G%%
]G%%
'G^%
R^HW
R^/W
]G5%
,"ANNgX`SXNNgX
&8;"ANNgX$'XNNgXf'^?"7=:3D
: A >"#>
;" AN NgX`SXN NgX
& 1>D: ?AAA" C
;"AN NgX$'XN NgXf'^ A4"@>B[?B64B6 >6?" >"4<?
B1? 9:;<
0
2
4
6
8
44b
LLE-TQ5 L LE-CB6 LLE-TQ5
- L LE-CB6
,"#4
!345G>>?`8?6D96
>?3<C;
D:
D:A
SS
9>I-H07d567;:7_9C=<\C_9C;
:G::7BD4
` A@80 =:3?"<" ?7?;"#44"4"#OQD9CD7D
A
)p#4B:4` ;"4?> ;\>:G 9<"D"#B9C<?B0=6OC:
#>"#;>E0A8`G4"V#>"#?"70AAAw#>"#B0=6 9<
OU>?[D8 94#A=#>w#>"#`B0=6 9<;?6
8"C9:D"#C?"7?7?;<
P7I-J0G:a`59B37BD\CaB>[7
`59BG B>[7
0
2
4
6
8
44b
LLEC12 LLEC12
-LLEX28
,"#4
D:
D:A
LLEX28
!345G>>?`8?6D96
>?3<C;
Sc
}`q "7E:": 9:>9CC4"@>>6;\="
: O" "04@ C# O" "<
A9C!l9!>#O"=#?@BCl4#
:
q`} "7C4": 9:>9CC4"@>>6;\="
:O""04@ C#O"0
G "# A9CE"?7? >#O"G
?"787?3?<?O"@9="?"74#
}`} "7C"6
9C C" ;V B# H > >6
A9C!="C7>@>
q`q
C ?: BC
O" >9: ;9
D7> >E6 =
0>#DI7?
.">9CC<D
X )p?7>6AG?"7B0=@=#>C?:4
C:;"BC="O6@B9C6AGB[?>1B9C<?O6@
>>1>E"@;" 9C"#B"0C9:>E"@
C:>E7;\
C6>6@>1 >E"@B"#8C74"@>>1?#>"B"74F
B\JZDEG$LSSK#4
AB"#>9C<B9C<? 9:;<=7
C748"?7??97034 G
?8C7
0#4B:4=7??!>#D><?B9C<?;?"9!?9780 9<D"#
C#O"?#>R$$cTGR$5^~
X
E"4"@>?9780 >"#?B"@D7> >E6?:;"BC8;"
A>E#4
J
E8 G;E "m4 `2'%%5KG
?7?>7?: AEQC:@ C"0 >E"4"
S^
>E9C>#O"A4>3=:3=#>C1?97?:O"@BC
.#>F:?97B!?aEEQ@ C
B!B7D97B9C3>"#?B"@D7>>E6>E4">E9C
>"#>6R$$cT
X "@> D>> B9C7? : 9C
<6 J
>;
_>> >>G4OE5L]&]SSK80D9CD7D ><" E9!?:#4
4EU 1> ?AAA
9 4"@> ?: #4
O
B0G>#>@O">9:BC8?"<A"<?,"BC
.X] ?"7 =: 3 D : A >"#>G ;" BC
]$1>D:?AAA" .>#>9<?@480=:3
D: >E "V A 4 J
G ;" BC.X]D : A8 ?:
?AAA" 0C;"BC]$4;"BCA8C7?7?>7?:
>B9C<??:
?"7=:3D:A?"
8 1>D:?AAA"
#D >#@G?7? >7?:@
># ?7? ?: ># @ :4 ;0 "[?4#> =: 3 ><" E ?AAA
R&$T
X _?4;> Ai 8 .OE= J$LL$K B! ; ?: #4 $
S5'c&B6 >@O" >9:G >#A8@ >#
E4"
>E9C4A>XEC:@>B"@'5X'S%G>#>?:?7?;"BCD7>>E6>EOH
>9 'X&G&44bG >E" = B"7 ?7? ;" A D7> >E6 >"#> CG >E OH SX
^44bRL%T
?9?"7B0=@D7>>E6C:F4"AC7CG>#B9C<?0
D7DA>E0;"#4CG9=#>F:?:>1@44?7>"
>9?"?: ?7? ?" >EH >E9C7?B"7?: >7?:
"9C7?G4"@>A09!=B\;D9CD7DAB6>H4E9!;"
SL
#4?"7=:3D:A>"#>81>D:?AAA" A">"C<D
A7#>3F4"?978>#D>#@>H?"7=:
3=74D7E0>1?#>A7>7=7?
I-H-94_9C567;>9B>>3C:G::7BD>U=3<4:
' M" +F %" .O
.&
& .
9=#>F:?""@??7?A"
>EG ?"# =:" 7> =: 3 D : A 8 D :
?AAA" ?:$&?:#4>E4">E9C4"0?EO";#>AA
"[?G '=6 >9=?#8"4" >E9C>E;dD>EGB" 9
>E6 CCG=#>F:B9C<?@C:O:]&
P7I-K0
#3>?`><
>?6D96>?3?3<C&:7/K:7BDMDDO
GU
:7
&96>?3?3<C&
'
5
S
^
P
MDDY
CO
'
5
S
^
P
MDDY
CO
]S'%$ $% $S ]% &% L^HK X $% $$ $5G& /^S/
_ $& ]% 5' &' L^K c $] $&G] $^ H^HK
c%
NgX 'L ]S 5^G] c5GS V^II X X X X
NgX
$]GS ']G] ]^GS ScGS S^JK X X X X
NgX` '% 'cG& ]^G& S% R^K X X X X
NgXf $LG] 'c ]$ S^GS S^KS X X X X
NgX' ]$G] ]5 5'G] c'G] V^WJ X X X X
NgX 'S ]%GS ]cG] &^G] R^HV X ' ] 5 W^II
NgX$c% $^ '5 5]G] c'G] V^WJ X X X X
NgX '5G& ]% 5]G& c'G& V^WL X X X X
`fX
N ''G& ]%G& ]S &L R^IR & c ^ L /^/I
`
X
N ''G] '^G] ]LG] &^ R^HK X $% $$ $' /^K
`f ]$ 5$ S%GS ^%GS /W^WR c $c $L '5 I
, $' $cG& '5 ]]GS J^H & $'G& $5G& '$ H^LH
$% $c '' ]$ I^SR c $' $SG] '$G& H^LV
E4" >E9C AG >6 ?" >E6 >"#>G >C #4
D7> >E6 >E OH >9 SG'&X$%G%c44b "0 >C ?7? ?: 0 ?!
D:?AAA" #G8B7?77AG>#>?:?7??:$
B9
8">1@4B0=""[?1>D7>>E6B9CE4">E9C
04 3 9: ;< ?7? ?:$
8"4<? B1? D :
AE#>97 ="`O =8 >=4 J$LL$K@>#
#43"6>#C:@>Aj_>=kJ$
?"7"B"@D:
A =" =74 H
7? >7? : B! >#
?"E4">E9CA8?AAA" 09>#>?:9!?:B9C<?
?""#>G >E"B"7?"7
4"@>?:?"7=:3 A4:44A9CC7]^G5M>E"
8"]%R'cT
?0D:A97EQ$
D7>>E6>"#>
C: @> B"@ =": ''%X 'S% . >91 @4 >E ?7? ?:
$
G ?7 >" B9 8" D" ?"7 @> B"@ "6 B\ A '&%#
c$
>E6=97;4A4"@>8@>BC7>H>9CB"#="7=3
8 >"# =748H D: I9: A7 @> B"@ D C 9C7
?97 >#D >" A ><" E 9! ?:$
?\ @> >1?
8C79C7?>
! >" =#> F: >9EG ?7? ?:G ;" #4 4"
JE?AEK 1> >": E >E6 8"#D >"4 D :
A>"#>91>D:?AAA"
7 d 5 6 7 : ; ; ; 9 8 \ C ;
: 7 / K : 7 BD
0
2
4
6
8
1 0
1 2
P C
3 6
2 0
1
S C
L E
C h
L E
T Q
L E
C B L E
X
L E
2 N L E
M
L E
1 7
0 L E
C
B X
T L
B T
T L
B X
V N A u
D A u
T
7
D D Y C
: ? A A A "
: A
9>I-I07d567:;;;98\C;:7/K:7BD
c'
@I-H0G::7BD5G>>?`>?6D96>?34:;8JC
@I-I0G::7BD5G>>?`>?6D96>?34:&8JC
' M"+F
3?978"=#>F:B\>1>EG?7>">6?"<9!?:#4
?"7=:3D:A>"#>9AD:?AAA" >#D>
=:3D:A?:4?9OQD9CD7DB\A9C<0A97
>9 =#> F: B\ >1 ?" ># >#> ?: ?7? ?: $
B9 8" >1
@4B0B7D 97 B9C6 ?"< B9C<? O""04
c]
GNgXGNgX
GNgX`GNgXfGNgX'GNgXGNgX$c%GNgXG`fX
NG`
X
NG`f.#>F:D>1?B\A9C<9
P7I-L0
CD;39><C96:BBDaG:
C
CD;3?9D96>?3<CDC:396:BBDMfO
]S'%$ NgX NgX
NgX` NgXf NgX' NgX NgX$c% NgX `fX
N `
X
N `f
$& LLG'^ LLG$5 LLG$& LLG'^ L^G^L L^Gc& LLG'^ LcGLS LLG%' L^G5& LLG'^ L^G']
$^ L^G'& L^G%L LcG&& L^Gc& L^G%L LcGLS L^Gc& LSGcc L^GS' LcG]' LLG%' LSG$$
'$ LSGL LSGS5 LSG'^ LcGc LcG%5 LSG&$ L^G%c L&G$L L&G5& L&GS$ L^Gc& L5G$]
'5 L5GS& L]Gc' L&G$' L&Gc' L&G]' L&G^& L&Gc' L]G%^ L5G5 L5 L5GSS L]GS$
'c L]G'S L$G^L L5 L]G%^ L&G$L L5 L&G$L L'GL& L]G5c L'G&5 L5G'c L]G'$
]% L$Gc' L%GLc L'G&] L'G&& L]G5c L'GL& L'GS^ L'G^$ L'GS^ L$G' L]G^c L'G5'
]] ^cG&S ^LG'& L$G' L%G]$ L%G55 L$G$ L$GcS ^LG$' L$GcS ^LG5c L'G^$ L$G]S
4A9C">C"?#G?"B#>97]]
?7??:#48V?">#D>E"4">E9C4?9=7
>"#>
`A@80=:3D:A8?AAA" ?:4"@> "##4B:4
` ;"4?> 4EU "8C74"@> "##4=7?
!?"79!?9780 9E0B"#>9C<9
?>"4?> G#44<?404G#44<?
)p4"@> "#A"?>"4?> G?7??97?:#>#?7?"7
=:3"7A":A8="7"7AC:497?B"@1>J?>G$L^cKC
?#D:A?:?7?A"?>"4?> ?9B9C09?7?
?97?!B!I7?B\B9C<?C:
DDG=:3D:
" B"@ ?:
9!P4=7? 9D"A"I; G ?AAA" GIA >H=" E"! E
R'^T
c5
#4 4 "74 ": >"@? AC7D ` ;"4?> ?"7 =:
D:?AAA" 89C>E"@ "8C7D:AG;"8@?"78>E"F>E"<
>E"8@?D7:"VGB[?O@>AD7:?7?E9>"J,IGWO >EG$LL&K
@??97>EB"#>9C<B#?"0="7=38H?7="
O6@">"">1D:>E4">E9C>1@4JgE= "8
?"@ 9<G$LL%K
#4 4"@? AC7D ?"4?> 8 ,>E"4?> >9C
D7:"V9C7>7A9!>7?D:?@481>>7?>E"
?7?A"#4D7:"V79>#?"8">D0D"A ??E;8
1>D:A"71>?97>EB"#>9C<?C?#D:?:
?7?9B9CE"!R&^T
9!<?#?:?7?"74#4=7?B!A4?""74#44<?
>EU?"74"@>8>E"F>E"E" 9 "##4
4EU D7> >E6 E#> >"#> >E A@ "V =" ?0 =9: >E 9
J
_>>&5c'^c5G
_>>&5cScL%KR'^T"@>8 "#A@80=:
3D:"V?:#44EU9 _5>7"?#>E4:
"V "0G & ?: $
A4 :4 $^X]5M >E"< A9C< ="# "VG
A4:4'LG]MG
A4:4&cGcMR'cTG
R$%STG R$]^T
E" "# 9!#44EUD:"VG
AA"B9C40#>GB[?O@>A
C:=:3D: A9 1>D:?AAA" G4C: E >E68"<?"
8@?>0I9:A7A@>E"?"@D#OQD9CD7D "<?
9!98@>?:A"#4G?7>" !OA@C:D0 R]$TGR^$T
I-I-7d567;:7E:7/. .) % .
c&
''6 /. .) % .
%%)&$
&"
A4"@>A"#4B:4?"7=:3D7:
"V >E" >9< "7 0 A" #4 D : "VG 9 ?7? ?:
B9CE" :
I#> O"@> # " "# B[? B64 9 1> >7? B"@ B#
?AAA" 8 ?"7 >6 >3 >E9C: >"#> C: @> B"@ =7 ?" J=": 5%%K 7?
?97]?:
"#?"<Q4B77=:
3D:A8?AAA" >E"B0=@=7?B6>H4?:>1?
C4R$5ST
'' 2P + & %" .O
.&
&. Q
B:
"
+ % 'NN6±N6
";"! 9>E6 CC?:]?:]S'%%G]S'%$G]S]$L
>E4">E9CEXA8EXG@B9CF:
P7I-R0!345G>>?`8=9U59C
.7@?: _9>E6 CC
>E4">E9C
AJ44K
_9>E6 CC
>E4">E9C
J44K
' '% $%
] '5 '%
5 ]% '&
]S'%%
K JW IH
' '$ &]S'%$
] '^ ^
cS
5 ]& $$
K JH /K
' '% &
] '& ^
5 ]' $'
]S]$L
K J/ /R
#>97&G":;;\?A"A8"4">E9C"?#B6F 7>
8"D:
.#>F:?">#X:]?:B0?"7=:3D:A>"#>
X : ]S'%% B"0 >C ?"7 =: 3 D :
?AAA" ?"C'?:?"A<
9>I-J0!345G>>?`8?6D96>?3<C:;;;98\C;
:7I:7/.) % .
=9U59C
!345G>>?`>?6D96>?3<C:;;;98\C;
:7I:7 /.) % .
\C9C>3GK
0
10
20
30
40
50
PC. 36200PC. 36201PC. 36319
7BD
mm
A>C>E4>
A>C>E4>A
E]S'%%
E4">E9CA
E]S'%%
E4">E9C
cc
@I-J0G::7"-5G>>?`>?6D96>?3<C;\C&8KC
B:
"
+&)
=NN6?NN6Q
6=64 >E@F: >3?9C" >1D: AG?7 >" >9:
@4 "#?C:@>B"@=7?"G=#>F:@B9CEHO>EO:]^
P7I-S0!345G>>?`8<:7I:7/.) % .
=9UKWW\CLWW
.7@
?:
_9>E6 CC>E
4">E9CAJ44K
_9>E6 CC>E
4">E9CJ44K
&%% S%% &%% S%%
]S'%% $ & & & &
E]S'%$
E4">E9CA
E]S'%$
E4">E9C
E]S]$L
E4">E9CA
E]S]$L
E4">E9C
c^
' 5' 55 '% '$
] S] S& ]^ 5%
J SK SK KW LV
$ & & & &
' 5% 5c L $]
] SS SS $^& 'L
]S'%$
J SK SK J/ JW
$ & & & &
' 5S 5& $5 $^
] S] S$ ]% ]^
]S]$L
J SK SK JS JK
!345G>>?`>?6D96>?3<C&\C;
\C9C>3GJ
0
10
20
30
40
50
60
70
80
90
PC.36200 PC.36201 PC.36319
7
mm
CMC
lignin
9>I-K0!345G>>?`8?6D96>?3<C:;;;98\C;:7I:7
/.) % .
=3<7KWW
@I-K07"-ILHW/5G>>?`>?6D96>?3<C;\C&MKWWO
cL
Z"! E A = >3@> B"@ >E" >CUG >3 >E9C:?:
?7??:
>1@4B0>6@89C>E"@ "8C7
C:@>B"@D"@>9C "#>1@4?97>E9C7?
BI7?@G "?7?>7?:>9C "#?C:@>B"@>#DCB"?7>
!345G>>?`>?6D96>?3<C&
\C;\C9C>3GJ
0
20
40
60
80
100
PC.36200 PC.36201 PC.36319
7
mm
CMC
lignin
9>I-L0!345G>>?`8?6D96>?3<C:;;;98\C;
:7I:7/.) % .
=3<7LWW
@I-L07"-ILHW/5G>>?`>?6D96>?3<C;\C&MLWWO
#4
"#?@>C:&%%8S%%B0?"7
=:3D:A>"#>CC:@>B"@D"#>975G>C#4B!
AB0"@DD>EB9C=1^&44 "8C7C:@>B"@D"B#>97&8V
?9 A B0 "@D >E 7 >" ?" C:
S%%B6B98"?97B\A9C<
^%
'' 1
.&
&
)
P7I-V0
CD;3?9D8><C96:BMfO
7 C &E@>3LWW
aA@4"
"#$b&
aA@4"
"#$b$%
aA@4"
"#$b'%
aA@4"
"#$b&
aA@4"
"#$b$%
aA@4"
"#$b'%
9B:3G /K^IKM>3<3G/WWf;O
"-ILHWW $& $%G'$ $%G&c $%G&^ $%G$L $%G'$ $%G$5
'& LGS^ $%G%& $%G5' LG&L LG%c LGS]
]& LG5] LG&S $%G%& ^GSS ^G^' LG]]
5& ^G'& ^G&] LG'5 cG^c ^G%$ LG$&
N:4
5&
R^/W
MJL^HKfY%O
L^SH
MJJ^JIfY%O
L^//
MIV^IJfY%O
R^JS
MJS^RIfY%O
R^IJ
MJR^SHfY%O
L^HW
MJW^IVfY%O
"-ILHW/ $& $%GcS $%G%$ $%G& LGLL LGS% LGc]
'& $%G$% LG]5 LG^c LG]& LG%L ^GL^
]& LG%] ^GL5 LG&^ ^GL& ^G^% ^GSc
5& ^Gcc ^G^L LG$& ^G&' ^GS% ^G]5
N:4
5&
L^KS
MJH^SLfY%O
L^JL
MJH^WSfY%O
L^HW
MJW^IVfY%O
L^SI
MJJ^JVfY%O
L^RK
MJI^VRfY%O
R^W/
MJK^LRfY%O
"-ILI/V $& LGc] $%G5& $%G^L LG^^ LGcc $%G%S
'& LG'L LG%$ $%G]% ^G]c LG$^ LG]$
]& ^G5$ ^GcS LG^& ^G$% ^G]% ^G5]
5& cGL] ^G5c ^GS$ cG'' cGc$ cG^L
N:4
5&
R^JH
MJS^IJfY%O
L^SS
MJJ^SHfY%O
L^RJ
MJI^VWfY%O
S^/I
MKH^VLfY%O
R^LJ
MJV^RRfY%O
R^JL
MJS^LWfY%O
_5&"?#8C7]>aA@4""#=7?G4A9C<A
>E"?C?#>:41>#>A]LG]5M8?"#>A&'GLSM`?:]S'%%G
]S'%$G]S]$L?"7=:3D:A>9CB9CG>E"B"7C:
?:]S]$L=:3?"7D0?"C'?:?"AA=
B9CC:S%%
'''1
.&
&
)
P7I-/W0
CD;3?9D8><C96:BMfO
7 C &E@>3LWW
aA@4"
aA@4"
aA@4"
aA@4"
aA@4"
aA@4"
^$
"#$b& "#$b$% "#$b'% "#$b& "#$b$% "#$b'%
9B:3G RJ^SKM>3<3G/WWf:;;;98O
"-ILHWW $& c$G^c c'G5^ SLG%5 c'G%c c'GcL c5G&]
'& S&G]5 S^G^L S^G^L c%G]' c'GSL c%GS]
]& S$G]^ ScG&& S^G]c S]Gc& c$G^$ c%G]'
5& &cG]^ S&G%^ S&Gc% S$GL% SSGL] S^G&^
AAA"
:4
5&
JMK
/R^JR
MHI^IJfYO
V^RR
M/I^WKfYO
V^/K
M/H^HHfYO
/H^VK
M/R^IWfYO
R^VH
M/W^KSfYO
L^HR
MS^ISfYO
"-ILHW/ $& S^G5c S^G^L c'Gc5 c]G^' c5G5] c]G]%
'& ScG'5 S^G]c SLG]& c$G]& c]G$% c%Gc]
]& SSG$$ S^G$c S^G&^ S&Gc% c$GSS SLGS%
5& S5G'S SSGc] ScG$L S&G%^ ScG]5 S^G^L
AAA"
:4
5&
JMK
/W^KV
M/J^/KfYO
S^/H
M/W^SKfYO
R^LL
M/W^HIfYO
V^RR
M/I^WKfYO
R^K/
M/W^WIfYO
K^VL
MR^VLfYO
"-ILI/V $& c$G&S c$G5& c$GcS c$GLc c'G$c c]GS$
'& SSGc] S^Gc^ c%G'' ScGcS c$GcS c$GLc
]& S5G'$ ScG&& SLG%L S&G'5 c%G$' SLG&%
5& S'G'% S5GSc S^G%S S]G&5 ScG&& S^Gc^
AAA"
:4
5&
JMK
/H^LK
M/L^VfYO
/W^/S
M/I^LfYO
L^RV
MV^WRfYO
//^I/
M/K^//fYO
R^IW
MV^RKfYO
L^WR
MS^//fYO
X
94VB"#?97OB0G 5&?#4"#48"?C?#>8C7?7?>aA@
=7?GC:4VOH>9C4A9C<?AAA" :40C4VB9C<?
"#?@>C:S%%)p@>B"@ "#?GP4D:AB9C<"E
0 >a A@ >7?B"@ A ?C?#> "8C7P4D:?AAA" ?"7 >E"@
C
X:]S'%% 5&"?#A4:44A9CE?C
._.